The MareCet team have a number of publications on marine mammals and other wildlife in Malaysia and further afield in peer-reviewed scientific journals and news bulletins. If you would like a copy of any of these, please send your request to

Verutes, G.M., Johnson, A.F., Caillat, M., Ponnampalam, L.S., Peter, C., Vu, L., Junchompoo, C., Lewison, R.L. and Hines, E.M. 2020. Using GIS and stakeholder involvement to innovate marine mammal bycatch risk assessment in data-limited fisheries. PLoS ONE 15(8): e0237835.

Vu, L.,  McGowen, M.R., Potter, C.W., Truong, A.T., Kuit, S.H.,  Abdel-Raheem, S.T. and Hines, E. 2020. New records of Fraser’s Dolphin (Lagenodelphis hosei) from the whale temples and fishing communities of Vietnam. Aquatic Mammals 46: 395 - 401. doi:

Hines, E., Ponnampalam, L.S., Junchompoo, C., Peter, C., Vu, L., Huynh, T., Caillat, M., Johnson, A.F., Minton, G., Lewison, R. and Verutes, G.M. 2020. Getting to the bottom of bycatch, a GIS-based toolbox to assess the risk of marine mammal bycatch. Endangered Species Research. doi:

Jackson‐Ricketts, J., Junchompoo, C., Hines, E.M., Hazen, E.L., Ponnampalam, L.S., Ilangakoon, A. and Monanunsap, S. 2020. Habitat modeling of Irrawaddy dolphins (Orcaella brevirostris) in the Eastern Gulf of Thailand. Ecology and Evolution 10: 2778 - 2792. doi:

Moura, A.E., Shreves, K., Pilot, M., Andrews, K.R., Moore, D.M., Kishida, T., Möller, L., Natoli, A., Gaspari, S., McGowen, M., Chen, I., Gray, H., Gore, M., Culloch, R.M., Kiani, M.S., Willson, M.S., Bulushi, A., Collins, T., Baldwin, R., Willson, A., Minton, G., Ponnampalam, L. and Hoelzel, A.R. 2020. Phylogenomics of the genus Tursiops and closely related Delphininae reveals extensive reticulation among lineages and provides inference about eco-evolutionary drivers. Molecular Phylogenetics and Evolution 146: 106756. doi:

Kuit, S.H., Ponnampalam, L.S., Ng, J.E., Chong, V.C. and Then, A.Y. 2019. Distribution and habitat characteristics of three sympatric cetacean species in the coastal waters of Matang, Perak, Peninsular Malaysia. Aquatic Conservation: Marine and Freshwater Ecosystems 29: 1681 - 1696. doi:

Lombardini, E.D., Haetrakul, T., Kuit, S.H. and Chansue, N. 2019. Gastric Braunina cordiformis and a review of helminth parasitism in the finless porpoise (Neophocaena phocaenoides). Brazilian Journal of Veterinary Pathology 12: 24 - 26.

Graham, I.M., Merchant, N. D., Farcas, A., Barton, T.R., Cheney, B., Bono, S. and Thompson, P.M. 2019. Harbour porpoise responses to pile-driving diminish over time. Royal Society Open Science 6: 190335.

Ponnampalam, L.S., Kuit, S.H. and Ng, J.E. 2018. Diversity and distribution of marine mammals in Tun Mustapha Park, Kudat, Sabah, during the Tun Mustapha Park Scientific Expedition 2017. Sabah Parks Nature Journal 11:35 – 46. Link to PDF here

Gray, H.W.I., Nishida, S., Welch, A.J., Moura, A.E., Tanabe, S., Kiani, M.S., Culloch, R., Möller, L., Natoli, A., Ponnampalam, L.S., Minton, G., Gore, M., Collins, T., Willson, A., Baldwin, R. and Hoelzel, A.R. 2018. Cryptic lineage differentiation among Indo-Pacific bottlenose dolphins (Tursiops aduncus) in the Northwest Indian Ocean. Molecular Phylogenetics and Evolution 122: 1-14.

Pilcher N.J., Adulyanukosol K., Das H., Davis P., Hines E., Kwan D., Marsh, H., Ponnampalam, L. and Reynolds, J. 2017. A low-cost solution for documenting distribution and abundance of endangered marine fauna and impacts from fisheries. PLoS ONE 12(12): e0190021.

Johnson, A.F., Caillat, M., Verutes, G.M., Peter, C., Junchumpoo, C., Vu, L., Ponnampalam, L.S., Lewison, R.L. and Hines, E.M. 2017. Poor fisheries struggle with U.S. import rule. Science 355(6329): 1031-1032. DOI: 10.1126/science.aam9153

Hoffman, J.M., Ponnampalam, L.S., Araújo-Wang, C., Wang, J.Y., Kuit, S.H. and Hung, S.K. 2016. Description of whistles of Irrawaddy dolphins (Orcaella brevirostris) from the waters of Matang, Peninsular Malaysia. Bioacoustics 26: 15 - 24. doi:

Ponnampalam, L. S. 2016. No danger in sight? An observation of sperm whales (Physeter macrocephalus) in marguerite formation off Muscat, Sultanate of Oman. 2016. Aquatic Mammals 42(2): 162-167. DOI 10.1578/AM.42.2.2016.162

Hoffman, J.M., Ponnampalam, L.S., Araújo, C.C., Wang, J.Y., Kuit, S.H. and Hung, S.K. 2015. Comparison of Indo-Pacific humpback dolphin (Sousa chinensis) whistles from two areas of western Peninsular Malaysia. Journal of the Acoustical Society of America 138(5): 2829-2835. doi:

Hines, E., Ponnampalam, L.S., Fairul Izmal, J.H., Whitty, T.S., Jackson-Ricketts, J., Kuit, S.H. and Acebes, J. 2015. Report of the Third Southeast Asian Marine Mammal Symposium (SEAMAM III). UNEP/CMS Secretariat Technical Series. Available online at:

Hines, E.M., Strindberg, S., Junchumpoo, C., Ponnampalam, L.S., Ilangakoon, A.D., Jackson-Ricketts, J. and Monanunsap, S. 2015. Line transect estimates of Irrawaddy dolphin abundance along the eastern Gulf Coast of Thailand. Frontiers in Marine Science 2:63. doi: 10.3389/fmars.2015.00063

Ichikawa, K., Akamatsu, T., Ponnampalam, L., Alfian, K. and Arai, N. 2015. Heterogeneous movement of dugongs between a vocal hotspot and a seagrass bed in Johor, Malaysia. Abstracts of the 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000), 15 March 2015, Kyoto, Japan.

Pomilla, C., Amaral, A. R., Collins, T., Minton, G., Findlay, K., Leslie, M. S., Ponnampalam, L., Baldwin, R. and Rosenbaum, H. 2014. The world’s most isolated and distinct whale population? Humpback whales of the Arabian Sea. PLoS ONE 9(12): e114162. doi: 10.1371/journal.pone.0114162

Ponnampalam, L.S., Fairul Izmal, J.H., Adulyanukosol, K., Ooi, J.L.S. and Reynolds III, J.E. 2015. Aligning conservation and research priorities for proactive species and habitat management: the case of dugongs Dugong dugon in Johor, Malaysia. Oryx 49: 743-749. doi: 10.1017/S0030605313001580  

Ponnampalam, L. S., Fairul Izmal, J. H., Ichikawa, K., Akamatsu, T., Kee Alfian, A. A., Wetzel D. and Reynolds III, J. 2014. A multi-pronged approach for overcoming knowledge barriers on the ecology and status of dugongs (Dugong dugon) in Johor, Malaysia: Towards critical habitat protection. Pages 101 – 112 in Wan Musthapa, W. F. Z., Nik Rosely, N. F. and Othman, A. S. (Eds). Johor Nature Heritage. School of Biological Sciences, Universiti Sains Malaysia, Penang, Malaysia.

Kuit, S. H., Ponnampalam, L. S., Fairul Izmal, J. H. and Chong, V. C. 2014. Cetacean research and a precautionary approach in developing dolphin-watching tourism in the coastal waters of the Matang mangroves. Proceedings of the Matang Mangrove Forest Management Conference 2014, Ipoh, Perak, Malaysia. pp. 27 – 39.

Ponnampalam, L. S. 2014. Medics and marine mammals – An unlikely but important connection for humanity’s survival. Malaysian Journal of Medical Sciences 21(2): 1- 3.

Ponnampalam, L. S. 2013. The dolphins of the Matang Mangroves. Pages 110 – 112 in Ariffin, R. and Nik Mustafa, N.M.S. (Eds.). A working plan for the Matang Mangrove Forest Reserve, Perak. 6th Revision. Perak State Forestry Department, Malaysia.

Ponnampalam, L. S., Hines, E. M., Monanunsap, S., Ilangakoon, A. D., Junchompoo, C., Adulyanukosol, K. and Morse, L. J. 2013. Behavioral observations of coastal Irrawaddy dolphins (Orcaella brevirostris) in Trat Province, Eastern Gulf of Thailand. Aquatic Mammals 39(4): 401 - 408. doi: 10.1578/AM.39.4.2013.401

Kimura, S., Ponnampalam, L., Akamatsu, T. and Yoda, K. 2013. Acoustic towing survey for marine mammals in Langkawi, Malaysia, with special reference to Indo-Pacific finless porpoise and Indo-Pacific humpback dolphin. Abstracts of the 9th International Symposium on SEASTAR2000 and Asian Bio-logging Science (The 13th SEASTAR2000 Workshop), 18 – 19 March 2013, Kyoto, Japan. pp. 13.

Ponnampalam, L. S., Collins, T., Minton, G., Schulz, I., Gray, H., Ormond, R. and Baldwin, R. 2012. Stomach contents of stranded cetaceans along the Sea of Oman and the Arabian Sea coasts of the Sultanate of Oman. Journal of the Marine Biological Association of the United Kingdom 92(8): 1699 - 1710. doi: 10.1017/S0025315411002104.

Ponnampalam, L. S. 2012. Opportunistic observations on the distribution of cetaceans in the Malaysian South China, Sulu and Sulawesi Seas and an updated checklist of marine mammals in Malaysia. The Raffles Bulletin of Zoology 60(1): 221 - 231. Link to PDF here

Ponnampalam, L. S. 2011. Dolphin watching in Muscat, Sultanate of Oman: Tourist perceptions and actual current practice. Tourism in Marine Environments 7(2): 81 – 93. Link to PDF here

Ponnampalam, L. S., F. I. Jamal Hisne, and C. R. Kaur. 2010. Report on the roundtable discussion on establishing a national marine mammal stranding network in Malaysia. MIMA: 3 August 2010. MIMA Bulletin 17: 38 – 42.

Gomez, L., Shepherd, C.R. and Khoo, M.S. 2020. The illegal bear bile trade in Sabah and Sarawak. Endangered Species Research 41: 279–287. DOI:

Gomez, L., Shepherd, C.R. and Morgan, J. 2019. The online trade of Sun Bears in Indonesia. TRAFFIC Bulletin Vol. 31 No. 2. Link to PDF here

Gomez, L. and Shepherd, C.R. 2019. Bearly on the radar: an analysis of bear seizure data in Indonesia. European Journal of Wildlife Research 65: 89. doi:

Gomez, L. 2019. An update on the bear bile trade in Peninsular Malaysia. TRAFFIC, Petaling Jaya, Malaysia. Link to PDF here

Gomez, L. and Krishnasamy, K. 2019. A rapid assessment of the trade in Saiga Antelope horn in Peninsular Malaysia. TRAFFIC Bulletin Vol. 31 No. 1. Link to PDF here

Gomez, L. and Krishnasamy, K. 2019. A rapid assessment on the trade of marine turtles in Indonesia, Malaysia and Vietnam. TRAFFIC, Petaling Jaya, Selangor, Malaysia.

Gomez, L., Shepherd, C.R. and Morgan, J. 2019. Improved legislation and stronger enforcement actions needed as the online otter trade in Indonesia continues. IUCN Otter Spec. Group Bull. 36(1). Link to PDF here

Leupen, B.T.C. and Gomez, L. 2019. The trading of the Orange-spotted Bulbul and Aceh Bulbul in Indonesia. BirdingASIA 32: 102–107. Link to PDF here

Janssen, J. and Gomez, L. 2019. Common Sun Skink, Eutropis multifasciata (Kuhl 1820) sold for Traditional Medicine in Indonesia and potential conservation implications. Ethnobiology and Conservation 8: 14. DOI: 10.15451/ec2019-11-8.14-1-8

Gomez, L. and Sy, E. 2018. Illegal pangolin trade in the Philippines. TRAFFIC Bulletin Vol. 20 No.1. Link to PDF here

Gomez, L. and Shepherd, C.R. 2018. Trade in bears in Lao PDR with observations from market surveys and seizure data. Global Ecology and Conservation 15: e00415. doi:

Gomez, L. and Bouhuys, J. 2018. Illegal otter trade in Southeast Asia. TRAFFIC, Petaling Jaya, Selangor, Malaysia. Link to PDF here

Kaur, R., Singh, S., Ramli, R., Hassan, H., Majinun, A., Ancrenaz, M., Ratag, M., Rauf, H. A., Jamal, J., Rauf, A. S. A., Yunus, M. D. M., Arbani, M. A., Asmar, K. and Amir, M. A. 2018. Observations at a nest of Helmeted Hornbill Rhinoplax vigil in Borneo, Malaysia. Forktail 34: 68 - 73. Link to PDF here

Shepherd, C.R. and Gomez, L. 2018. Trade and conservation efforts involving the Sumatran Laughing Thrush in Indonesia. Journal of Indonesian Natural History Vol 6 No 2. Link to PDF here

Livingstone, E., Gomez, L. and Bouhuys, J. 2018. A review of bear farming and bear trade in Lao PDR. Global Ecology and Conservation 13(2018) e00380. doi:

Gomez, L. Leupen, B.T.C., Krishnasamy, K. and Heinrich, S. 2017. Scaly nexus: mapping Indonesian pangolin seizures (2010-2015). TRAFFIC, Petaling Jaya, Malaysia. Link to PDF here

Leupen, B.T.C, Gomez, L. and Shepherd, C.R. 2017. Recent observations of the illegal trade in serows in Lao PDR. TRAFFIC Bulletin 29: 37 - 40.  Link to PDF here

Or, O.C., Gomez, L. and Lau, C.F. 2017. Recent reports of seizures and poaching of sun bears in Malaysia. International Bear News 26 (2): 17 - 18

Gomez, L. and Bouhuys, J. 2017. Recent seizures of otters in Southeast Asia. IUCN Otter Spec. Group Bull. 34(2). Link to PDF here

Willcox, D., Nguyen, D.T. and Gomez, L. 2016. An assessment of trade: in bear bile and gall bladder in Viet Nam. TRAFFIC, Petaling Jaya, Selangor, Malaysia. Link to PDF here

Gomez, L., Leupen, B T.C., Theng, M., Fernandez, K., and Savage, M. 2016. Illegal otter trade: an analysis of seizures in selected Asian countries (1980-2015). TRAFFIC. Petaling Jaya, Selangor, Malaysia. Link to PDF here

Gomez, L., Leupen, B.T.C., Heinrich, S. 2016. Observations of the illegal pangolin trade in Lao PDR, 2016. TRAFFIC Southeast Asia Regional Office, Petaling Jaya, Selangor, Malaysia. Link to PDF here

Gomez, L., Leupen, B.T.C. and Tian, K.H. 2016. The trade of African pangolins to Asia: a brief case study of pangolin shipments from Nigeria. TRAFFIC Bulletin 28(1): 3 - 5. Link to PDF here

Kaur, R, Singh, S. and Ahmad, A. H. 2015. The breeding biology of the Great Hornbill Buceros bicornis, Rhinoceros Hornbill Buceros rhinoceros and Helmeted Hornbill Rhinoplax vigil in the Temengor Forest Reserve, Perak, Malaysia. Malayan Nature Journal 67: 242 - 259. Link to Abstract here

Tsai, S., Kuit, V., Lin, Z. G. and Lin, C.  2014. Application of a functional marker for the effect of cryoprotectant agents on gorgonian coral (Junceela juncea and J. fragilis) sperm sacs. Cryo Letters 35: 1 - 7.

Kaur, R., Ong, T., Lim, K.C. and Yeap, C.A. 2011. A survey on mass movements of the vulnerable Plain-Pouched Hornbill in the Belum-Temengor Forest Complex, Peninsular Malaysia. The Raffles Bulletin Of Zoology Supplement No. 24: 171 - 176. Link to PDF here

Yule, C.M. and Gomez, L.G. 2008. Leaf litter decomposition in a tropical peat swamp forest in Peninsular Malaysia. Wetlands Ecology and Management 17: 231 - 241. doi: